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All images generated by Dream-AI. TEAL Lab Members are highlighted in bold.

IMG_1569 Shrieking Skull Biker Tattoo.JPG

Wilson T, Szpak P. 2023. Comparing the performance of demineralization agents (HCl and EDTA) for stable isotope analysis of bone collagen with implications for quality control criteria and collagen yield. International Journal of Osteoarchaeology. doi:10.1002/oa.3222.



Dolphin AE, Teeter MA, Szpak P, 2023. The role of status, diets, and mobility in understanding the impacts of urbanization in early medieval Bergen, Norway (St. Mary's Church): Insights from stable isotope analyses. International Journal of Osteoarchaeology. doi: 10.1002/oa.3216. 



meps swordfish IMG_8991.JPG

Elliott Smith EA, Szpak P, Braje TJ, Newsom B, Rick TC, 2023. Pre-industrial ecology and foraging behavior of swordfish Xiphias gladius in the eastern North Pacific. Marine Ecology Progress Series 711, 129-134.[DOWNLOAD .pdf]




Szpak P. 2022. Why Zooarchaeology Needs Stable Isotope Analysis. In: Sharpe AE, and Krigbaum J, editors. Applications of Isotope Research in Zooarchaeology. Gainesville: University Press of Florida. p 248-270. 



Wilson T, Szpak P. 2022. Examining the use of ethylenediaminetetraacetic acid for humic extraction of ancient bone. American Journal of Biological Anthropology 179, 148-157. doi:10.1002/ajpa.24577. 



Guiry EJ, Orchard TJ, Needs-Howarth S, Szpak P, 2022. Freshwater wetland–driven variation in sulfur isotope compositions: Implications for human paleodiet and ecological research. Frontiers in Ecology and Evolution 10. [DOWNLOAD .pdf]


Louis M, Routledge J, Heide-Jørgensen MP, Szpak P, Lorenzen ED, 2022. Sex and size matter: foraging ecology of offshore harbour porpoises in waters around Greenland. Marine Biology 169, 140. doi:10.1007/s00227-022-04123-x.


IMG_1942 Purple Scientist.JPG

Rey-Iglesia A, Wilson T, Routledge J, Skovrind M, Garde E, Heide-Jørgensen MP, Szpak P, Lorenzen ED, 2022. Combining δ13C and δ15N from bone and dentine in marine mammal palaeoecological research: insights from toothed whales. Isotopes in Environmental and Health Studies, 1-12. doi:10.1080/10256016.2022.2145285.





Hyland C, Millaire J-F, Szpak P, 2021. Migration and maize in the Virú Valley: Understanding life histories through multi-tissue carbon, nitrogen, sulfur, and strontium isotope analyses. American Journal of Physical Anthropology 176, 21-35. doi:10.1002/ajpa.24271.



Guiry E, Orchard TJ, Needs-Howarth S, Szpak P, 2021. Isotopic Evidence for Garden Hunting and Resource Depression in the Late Woodland of Northeastern North America. American Antiquity 86, 90-110. doi:10.1017/aaq.2020.86. [DOWNLOAD .pdf]


Keighley X, Bro-Jørgensen MH, Ahlgren H, Szpak P, Ciucani MM, Sánchez Barreiro F, Howse L, Gotfredsen AB, Glykou A, Jordan P, Lidén K, Tange Olsen M, 2021. Predicting sample success for large-scale ancient DNA studies on marine mammals. Molecular Ecology Resources 21, 1149-1166. doi:10.1111/1755-0998.13331. [DOWNLOAD .pdf]


McCormack J, Szpak P, Bourgon N, Richards M, Hyland C, Méjean P, Hublin J-J, Jaouen K, 2021. Zinc isotopes from archaeological bones provide reliable tropic level information for marine mammals. Communications Biology 4, 683. doi:10.1038/s42003-021-02212-z. [DOWNLOAD .pdf]


Hyland C, Scott MB, Routledge J, Szpak P, 2022. Stable Carbon and Nitrogen Isotope Variability of Bone Collagen to Determine the Number of Isotopically Distinct Specimens. Journal of Archaeological Method and Theory 29, 666-686. doi:10.1007/s10816-021-09533-7. 


Cheung C, Szpak P, 2022. Interpreting Past Human Diets Using Stable Isotope Mixing Models—Best Practices for Data Acquisition. Journal of Archaeological Method and Theory 29, 138-161. doi:10.1007/s10816-021-09514-w.


Losey RJ, Nomokonova T, Guiry E, Fleming LS, Garvie-Lok SJ, Waters-Rist AL, Bieraugle M, Szpak P, et al., The evolution of dog diet and foraging: Insights from archaeological canids in Siberia. Science Advances 8, eabo6493. doi:10.1126/sciadv.abo6493. 


Pilaar Birch SE, and Szpak P. 2022. Current developments and future directions in archaeological science. Proceedings of the National Academy of Sciences 119(43):e2212490119. 


Vicari D, Lorenzen ED, Skovrind M, Szpak P, Louis M, Olsen MT, Brown RP, Lambert O, Bianucci G, Sabin RC, Meloro C, 2022. Skull ecomorphological variation of narwhals (Monodon monoceros, Linnaeus 1758) and belugas (Delphinapterus leucas, Pallas 1776) reveals phenotype of their hybrids. PLOS ONE 17, e0273122. doi:10.1371/journal.pone.027312. [DOWNLOAD .pdf]

Guiry EJ, Kennedy JR, O’Connell MT, Gray DR, Grant C, Szpak P, 2021. Early evidence for historical overfishing in the Gulf of Mexico. Science Advances 7, eabh2525. doi:10.1126/sciadv.abh2525. [DOWNLOAD .pdf]

Guiry EJ, Szpak P, 2021. Improved quality control criteria for stable carbon and nitrogen isotope measurements of ancient bone collagen. Journal of Archaeological Science 132, 105416. doi:10.1016/j.jas.2021.105416.[DOWNLOAD .pdf]

Louis M, Skovrind M, Garde E, Heide-Jørgensen MP, Szpak P, Lorenzen ED. 2021. Population-specific sex and size variation in long-term foraging ecology of belugas and narwhals. Royal Society Open Science 7, 202226. doi: 10.1098/rsos.202226. [DOWNLOAD .pdf]

Rey-Iglesia A, Lister AM, Stuart AJ, Bocherens H, Szpak P, Willerslev E, Lorenzen ED, 2021. Late Pleistocene paleoecology and phylogeography of woolly rhinoceroses. Quaternary Science Reviews 263, 106993. doi:10.1016/j.quascirev.2021.106993. [DOWNLOAD .pdf]



Szpak P, Buckley M, 2020. Sulfur isotopes (δ34S) in Arctic marine mammals: indicators of benthic vs. pelagic foraging. Marine Ecology Progress Series 653, 205-216. doi:10.3354/meps13493. [DOWNLOAD .pdf]


Szpak P, Chiou KL, 2020. A comparison of nitrogen isotope compositions of charred and desiccated botanical remains from northern Peru. Vegetation History and Archaeobotany 29, 527-538. doi:10.1007/s00334-019-00761-2.[DOWNLOAD .pdf]


Szpak P, Millaire J-F, Chapdelaine C, White CD, Longstaffe FJ, 2020. An Integrated isotopic study of Early Intermediate Period camelid husbandry in the Santa Valley, Perú. Environmental Archaeology, 25, 279-295. doi:10.1080/14614103.2019.1583302. [DOWNLOAD .pdf]


Szpak P, Valenzuela D, 2020. Camelid husbandry in the Atacama Desert? A stable isotope study of camelid bone collagen and textiles from the Lluta and Camarones Valleys, northern Chile. PLOS One 15, e0228332. doi:10.1371/journal.pone.0228332. [DOWNLOAD .pdf]


Szpak P, Julien M-H, Royle TCA, Savelle JM, Yang DY, Richards MP, 2020. Sexual differences in the foraging ecology of 19th century beluga whales (Delphinapterus leucas) from the Canadian High Arctic. Marine Mammal Science 36, 451-471. doi:10.1111/mms.12655. [DOWNLOAD .pdf]


Cheung C, Szpak P, 2020. Interpreting Past Human Diets Using Stable Isotope Mixing Models. Journal of Archaeological Method and Theory. doi:10.1007/s10816-020-09492-5.



Guiry EJ, Buckley M, Orchard TJ, Hawkins AL, Needs-Howarth S, Holm E, Szpak P, 2020. Deforestation caused abrupt shift in Great Lakes nitrogen cycle. Limnology and Oceanography 65, 1921-1935. doi:10.1002/lno.11428.



Guiry E, Royle TCA, Matson RG, Ward H, Weir T, Waber N, Brown TJ, Hunt BPV, Price MHH, Finney BP, Kaeriyama M, Qin Y, Yang DY, Szpak P. 2020 Differentiating salmonid migratory ecotypes through stable isotope analysis of collagen: Archaeological and ecological applications. PLOS One 15:e0232180. doi: 10.1371/journal.pone.0232180. [DOWNLOAD .pdf]


Guiry E, Royle TCA, Orchard TJ, Needs-Howarth S, Yang DY, Szpak P, 2020. Evidence for freshwater residency among Lake Ontario Atlantic salmon (Salmo salar) spawning in New York. Journal of Great Lakes Research. 46, 1036-1043. doi:10.1016/j.jglr.2020.05.009. [DOWNLOAD .pdf]


Guiry E, Szpak P, 2020. Quality Control for Modern Bone Collagen Stable Carbon and Nitrogen Isotope Measurements. Methods in Ecology and Evolution 11, 1049-1060. doi:10.1111/2041-210X.13433. [DOWNLOAD .pdf]


Guiry E, Szpak P, 2020. Seaweed-eating sheep show that δ34S evidence for marine diets can be fully masked by sea spray effects. Rapid Communications in Mass Spectrometry 34, e8868. doi:10.1002/rcm.8868.



Losey RJ, Guiry E, Nomokonova T, Gusev AV, Szpak P, 2020. Storing fish?: a dog’s isotopic biography provides insight into Iron Age food preservation strategies in the Russian Arctic. Archaeological and Anthropological Sciences. doi:10.1007/s12520-020-01166-3. [DOWNLOAD .pdf]



Szpak P, Longstaffe FJ, Macdonald R, Millaire J-F, White CD, Richards MP, 2019. Plant sulfur isotopic compositions are altered by marine fertilizers. Archaeological and Anthropological Sciences 11, 2989-2999. doi:10.1007/s12520-018-0716-5. [DOWNLOAD .pdf]


Szpak P, Savelle JM, Conolly J, Richards MP, 2019. Variation in Late Holocene Marine Environments in the Canadian Arctic Archipelago: Evidence from Ringed Seal Bone Collagen Stable Isotope Compositions. Quaternary Science Reviews 211, 136-155. doi:10.1016/j.quascirev.2019.03.016. [DOWNLOAD .pdf]


Delsuc F, Kuch M, Gibb GC, Karpinski E, Hackenberger D, Szpak P, Martínez JG, Mead JI, McDonald HG, MacPhee RDE, Billet G, Hautier L, Poinar HN, 2019. Ancient Mitogenomes Reveal the Evolutionary History and Biogeography of Sloths. Current Biology 29, 2031-2042.e2036. doi:10.1016/j.cub.2019.05.043. [DOWNLOAD .pdf]


Rey-Iglesia A, García-Vázquez A, Treadaway EC, van der Plicht J, Baryshnikov GF, Szpak P, Bocherens H, Boeskorov GG, Lorenzen ED, 2019. Evolutionary history and palaeoecology of brown bear in North-East Siberia re-examined using ancient DNA and stable isotopes from skeletal remains. Scientific Reports 9, 4462. doi:10.1038/s41598-019-40168-7. [DOWNLOAD .pdf]


Skovrind M, Castruita JAS, Haile J, Treadaway EC, Gopalakrishnan S, Westbury MV, Heide-Jørgensen MP, Szpak P, Lorenzen ED, 2019. Hybridization between two high Arctic cetaceans confirmed by genomic analysis. Scientific Reports 9, 7729. doi:10.1038/s41598-019-44038-0.

[DOWLOAD .pdf]


Dyke AS, Savelle JM, Szpak P, Southon JR, Howse L, Desrosiers PM, Kotar K, 2019. An Assessment of Marine Reservoir Corrections for Radiocarbon Dates on Walrus from the Foxe Basin Region of Arctic Canada. Radiocarbon 61, 67-81. doi:10.1017/RDC.2018.50. [DOWNLOAD .pdf]


Santoro CM, Gayo EM, ..., Szpak P, 2019. From the Pacific to the tropical forests: Networks of social interaction in the Atacama Desert, Late in the Pleistocene. Chungara Revista de Antropología Chilena 51, 5-25. doi:10.4067/S0717-73562019005000602.




Szpak P, Buckley M, Darwent CM, Richards MP, 2018. Long-term ecological changes in marine mammals driven by recent warming in northwestern Alaska. Global Change Biology 24, 490-503. doi:10.1111/gcb.13880. [DOWNLOAD .pdf]


Szpak P, Millaire JF, White CD, Donnan CB, Longstaffe FJ, 2018. Stable Isotope Sourcing of Wool from Textiles at Pacatnamú. Archaeometry 60, 612-627. doi:10.1111/arcm.12342. [DOWNLOAD .pdf]


Craine JM, Elmore AJ, ..., Szpak P, Templer PH, Van der Colff D, Werner C, Xu X, Yang Y, Yu G, Zmudczyńska-Skarbek K, 2018. Isotopic evidence for oligotrophication of terrestrial ecosystems. Nature Ecology & Evolution 2, 1735-1744. doi:10.1038/s41559-018-0694-0. [DOWNLOAD .pdf]


Delsuc F, Kuch M, Gibb GC, Hughes J, Szpak P, Southon J, Enk J, Duggan AT, Poinar HN, 2018. Resolving the phylogenetic position of Darwin's extinct ground sloth (Mylodon darwinii) using mitogenomic and nuclear exon data. Proceedings of the Royal Society B: Biological Sciences 285, 20180214. doi:10.1098/rspb.2018.0214. [DOWNLOAD .pdf]


Guiry E, Beglane F, Szpak P, Schulting R, McCormick F, Richards MP, 2018. Anthropogenic changes to the Holocene nitrogen cycle in Ireland. Science Advances 4, eaas9383. doi:10.1126/sciadv.aas9383. [DOWNLOAD .pdf]



Szpak P, Krippner K, Richards MP, 2017. Effects of Sodium Hydroxide Treatment and Ultrafiltration on the Removal of Humic Contaminants from Archaeological Bone. International Journal of Osteoarchaeology 27, 1070-1077. doi:10.1002/oa.2630.



Szpak P, Metcalfe JZ, Macdonald RA, 2017. Best Practices for Calibrating and Reporting Stable Isotope Measurements in Archaeology. Journal of Archaeological Science: Reports 13, 609-616. doi:10.1016/j.jasrep.2017.05.007. [DOWNLOAD .pdf] Appendices: [Appendix A] [Appendix B] [Appendix C] [Appendix D] [Appendix E] [Appendix F] [Appendix G]


Braje TJ, Rick TC, Szpak P, Newsome SD, McCain JM, Elliott Smith EA, Glassow M, Hamilton SL, 2017. Historical ecology and the conservation of large, hermaphroditic fishes in Pacific Coast kelp forest ecosystems. Science Advances 3, e1601759. doi:10.1126/sciadv.1601759. [DOWNLOAD .pdf] 


Guiry E, Szpak P, Richards MP, 2017. Isotopic analyses reveal geographical and socioeconomic patterns in historical domestic animal trade between predominantly wheat- and maize-growing agricultural regions in eastern North America. American Antiquity 82, 341-352. doi:10.1017/aaq.2016.34. [DOWNLOAD .pdf] 


Carlson RL, Szpak P, Richards MP, 2017. The Pender Canal Site and Beginnings of the Northwest Coast Cultural System. Canadian Journal Of Archaeology 41, 1-29. 




Szpak P, Chicoine D, Millaire J-F, White CD, Parry R, Longstaffe FJ, 2016. Early Horizon camelid management practices in the Nepeña Valley, north-central coast of Peru. Environmental Archaeology 21, 230-245.



Szpak P, Millaire JF, White CD, Bourget S, Longstaffe FJ, 2016. Life Histories of Sacrificed Camelids from Huancaco (Virú Valley). In: Klaus HD, Toyne JM. (Eds.), Reconstructing Sacrifice on the North Coast of Peru: Archaeological Studies of Ritual Violence in the Ancient Andes, University of Texas Press, Austin, pp. 319-341.​

[DOWNLOAD .pdf] 


Enk J, Devault A, Widga C, Saunders J, Szpak P, ..., Poinar H, 2016. Mammuthus Population Dynamics in Late Pleistocene North America: Divergence, Phylogeography and Introgression. Frontiers in Ecology and Evolution 4, 42. doi:10.3389/fevo.2016.00042. [DOWNLOAD .pdf]


Guiry EJ, Needs-Howarth S, Friedland KD, Hawkins AL, Szpak P, Macdonald R, Courtemanche M, Holm E, Richards MP, 2016. Lake Ontario salmon (Salmo salar) were not migratory: A long-standing historical debate solved through stable isotope analysis. Scientific Reports 6, 36249. doi:10.1038/srep36249. 



Guiry EJ, Szpak P, Richards MP, 2016. Effects of lipid extraction and ultrafiltration on stable carbon and nitrogen isotopic compositions of fish bone collagen. Rapid Communications in Mass Spectrometry 30, 1591-1600. doi:10.1002/rcm.7590.



Jaouen K, Szpak P, Richards MP, 2016. Zinc Isotope Ratios as Indicators of Diet and Trophic Level in Arctic Marine Mammals. PLoS ONE 11, e0152299. doi:10.1371/journal.pone.0152299. [DOWNLOAD .pdf]



Szpak P, 2015. Manures and Other Fertilizers, Identification and Analysis. In: Beaudry MC, Metheny K. (Eds.), The Archaeology of Food: An Encyclopedia, AltaMira Press, Walnut Creek, CA, pp. 300-301.


Szpak P, Millaire J-F, White CD, Lau GF, Surette F, Longstaffe FJ, 2015. Origins of Prehispanic Camelid Wool Textiles from the North and Central Coasts of Peru traced by Carbon and Nitrogen Isotopic Analyses. Current Anthropology 56, 449-459. doi:10.1086 /680873.



Orchard TJ, Szpak P, 2015. Zooarchaeological and Isotopic Insights into Locally Variable Economic Patterns: A Case Study from Late Holocene Southern Haida Gwaii, British Columbia. BC Studies 187, 107-147. 




Szpak P, 2014. Complexities of Nitrogen Isotope Biogeochemistry in Plant-Soil Systems: Implications for the Study of Ancient Agricultural and Animal Management Practices. Frontiers in Plant Science 5, 288. doi:10.3389/fpls.2014.00288 [DOWNLOAD .pdf]


Szpak P, Longstaffe FJ, Millaire J-F, White CD, 2014. Large variation in nitrogen isotopic composition of a fertilized legume. Journal of Archaeological Science 45, 72-79.  doi:10.1016/j.jas.2014.02.007 [DOWNLOAD .pdf]


Szpak P, Millaire J-F, White CD, Longstaffe FJ, 2014. Small scale camelid husbandry on the north coast of Peru (Virú Valley): Insight from stable isotope analysis. Journal of Anthropological Archaeology 36, 110-129. doi:10.1016/j.jaa.2014.08.005 [DOWNLOAD .pdf]

2013 and earlier


Szpak P, Orchard TJ, Salomon AK, Gröcke DR, 2013. Regional ecological variability and impact of the maritime fur trade on nearshore ecosystems in southern Haida Gwaii (British Columbia, Canada): evidence from stable isotope analysis of rockfish (Sebastes spp.) bone collagen. Archaeological and Anthropological Sciences 5, 159-182. 



Szpak P, White CD, Longstaffe FJ, Millaire J-F, Vásquez Sánchez VF, 2013. Carbon and Nitrogen Isotopic Survey of Northern Peruvian Plants: Baselines for Paleodietary and Paleoecological Studies. PLoS One 8, e53763. doi:10.1371/journal.pone.0053763 [DOWNLOAD .pdf]


Szpak P, Longstaffe FJ, Millaire J-F, White CD, 2012. Stable Isotope Biogeochemistry of Seabird Guano Fertilization: Results from Growth Chamber Studies with Maize (Zea mays). PLoS One 7, e33741. doi:10.1371/journal.pone.0033741 [DOWNLOAD .pdf]


Szpak P, Millaire J-F, White CD, Longstaffe FJ, 2012. Influence of seabird guano and camelid dung fertilization on the nitrogen isotopic composition of field-grown maize (Zea mays). Journal of Archaeological Science 39, 3721-3740. doi:10.1016/j.jas.2012.06.035 [DOWNLOAD .pdf​]


Szpak P, Orchard TJ, McKechnie I, Gröcke DR, 2012. Historical ecology of late Holocene sea otters (Enhydra lutris) from northern British Columbia: isotopic and zooarchaeological perspectives. Journal of Archaeological Science 39, 1553-1571. doi:10.1016/j.jas.2011.12.006 [DOWNLOAD .pdf​]


Szpak P, 2011. Fish bone chemistry and ultrastructure: implications for taphonomy and stable isotope analysis. Journal of Archaeological Science 38, 3358-3372. doi:10.1016/j.jas.2011.07.022 [DOWNLOAD .pdf​]


Orchard TJ, Szpak P, 2011. Identification of salmon species from archaeological remains on the northwest coast. In: Moss M, Cannon A. (Eds.), The Archaeology of North Pacific Fisheries, University of Alaska Press, Fairbanks, pp. 17-29.



Szpak P, Gröcke DR, Debruyne R, MacPhee RDE, Guthrie RD, Froese D, Zazula GD, Patterson WP, Poinar HN, 2010. Regional differences in bone collagen δ13C and δ15N of Pleistocene mammoths: Implications for paleoecology of the mammoth steppe. Palaeogeography, Palaeoclimatology, Palaeoecology 286, 88-96. [DOWNLOAD .pdf​]


Szpak P, Orchard TJ, Gröcke DR, 2009. A Late Holocene vertebrate food web from southern Haida Gwaii (Queen Charlotte Islands, British Columbia). Journal of Archaeological Science 36, 2734-2741. doi:10.1016/j.jas.2009.08.013 [DOWNLOAD .pdf​]


Debruyne R, Chu G, King CE, Bos K, Kuch M, Schwarz C, Szpak P, ..., Poinar HN, 2008. Out of America: ancient DNA evidence for a New World origin of Late Quaternary woolly mammoths. Current Biology 18, 1320-1326. doi:10.1016/j.cub.2008.07.061. [DOWNLOAD .pdf​]

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