© 2019 by Paul Szpak. 

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Articles in Refereed Journals


  • Szpak, P., Longstaffe, F.J., Macdonald, R., Millaire, J.-F., White, C.D., Richards, M.P., 2019. Plant sulfur isotopic compositions are altered by marine fertilizers. Archaeological and Anthropological Sciences 11, 2989-2999. doi:10.1007/s12520-018-0716-5 [DOWNLOAD .pdf]

  • Szpak, P., Millaire, J.-F., Chapdelaine, C., White, C.D., Longstaffe, F.J., 2019. An Integrated isotopic study of Early Intermediate Period camelid husbandry in the Santa Valley, Perú. Environmental Archaeology, 1-17. doi:10.1080/14614103.2019.1583302 [DOWNLOAD .pdf]

  • Szpak, P., Savelle, J.M., Conolly, J., Richards, M.P., 2019. Variation in Late Holocene Marine Environments in the Canadian Arctic Archipelago: Evidence from Ringed Seal Bone Collagen Stable Isotope Compositions. Quaternary Science Reviews 211, 136-155. doi:10.1016/j.quascirev.2019.03.016. [DOWNLOAD .pdf]

  • Delsuc, F., Kuch, M., Gibb, G.C., Karpinski, E., Hackenberger, D., Szpak, P., Martínez, J.G., Mead, J.I., McDonald, H.G., MacPhee, R.D.E., Billet, G., Hautier, L., Poinar, H.N., 2019. Ancient Mitogenomes Reveal the Evolutionary History and Biogeography of Sloths. Current Biology. doi:10.1016/j.cub.2019.05.043. [DOWNLOAD .pdf]

  • Dyke, A.S., Savelle, J.M., Szpak, P., Southon, J.R., Howse, L., Desrosiers, P.M., Kotar, K., 2019. An assessment of marine reservoir corrections for radiocarbon dates on walrus from the Foxe Basin region of Arctic Canada. Radiocarbon 61, 67-81. doi:10.1017/RDC.2018.50. [DOWNLOAD .pdf]

  • Rey-Iglesia, A., García-Vázquez, A., Treadaway, E.C., van der Plicht, J., Baryshnikov, G.F., Szpak, P., Bocherens, H., Boeskorov, G.G., Lorenzen, E.D., 2019. Evolutionary history and palaeoecology of brown bear in North-East Siberia re-examined using ancient DNA and stable isotopes from skeletal remains. Scientific Reports 9, 4462. doi:10.1038/s41598-019-40168-7. [DOWNLOAD .pdf]

  • Santoro, C.M., Gayo, E.M., Capriles, J.M., Rivadeneira, M.M., Herrera, K.A., Mandakovic, V., Rallo, M., Rech, J.A., Cases, B., Briones, L., Olguín, L., Valenzuela, D., Borrero, L.A., Ugalde, P.C., Rothhammer, F., Latorre, C., Szpak, P., 2019. From the Pacific to the tropical forests: Networks of social interaction in the Atacama Desert, Late in the Pleistocene. Chungara Revista de Antropología Chilena 51, 5-25. doi:10.4067/S0717-73562019005000602. [DOWNLOAD .pdf]

  • Skovrind, M., Castruita, J.A.S., Haile, J., Treadaway, E.C., Gopalakrishnan, S., Westbury, M.V., Heide-Jørgensen, M.P., Szpak, P., Lorenzen, E.D., 2019. Hybridization between two high Arctic cetaceans confirmed by genomic analysis. Scientific Reports 9, 7729. doi:10.1038/s41598-019-44038-0. [DOWLOAD .pdf]


  • Szpak, P., Buckley, M., Darwent, C.M., Richards, M.P., 2018. Long-term ecological changes in marine mammals driven by recent warming in northwestern Alaska. Global Change Biology 24, 490-503. doi:10.1111/gcb.13880 [DOWNLOAD .pdf]

  • Szpak, P., Millaire, J.F., White, C.D., Donnan, C.B., Longstaffe, F.J., 2018. Stable Isotope Sourcing of Wool from Textiles at Pacatnamú. Archaeometry 60, 612-627. doi:10.1111/arcm.12342 [DOWNLOAD .pdf]

  • Craine, J.M., Elmore, A.J., Wang, L., Aranibar, J., Bauters, M., Boeckx, P., Crowley, B.E., Dawes, M.A., Delzon, S., Fajardo, A., Fang, Y., Fujiyoshi, L., Gray, A., Guerrieri, R., Gundale, M.J., Hawke, D.J., Hietz, P., Jonard, M., Kearsley, E., Kenzo, T., Makarov, M., Marañón-Jiménez, S., McGlynn, T.P., McNeil, B.E., Mosher, S.G., Nelson, D.M., Peri, P.L., Roggy, J.C., Sanders-DeMott, R., Song, M., Szpak, P., Templer, P.H., Van der Colff, D., Werner, C., Xu, X., Yang, Y., Yu, G., Zmudczyńska-Skarbek, K., 2018. Isotopic evidence for oligotrophication of terrestrial ecosystems. Nature Ecology & Evolution 2, 1735-1744. doi:10.1038/s41559-018-0694-0. [DOWNLOAD .pdf]

  • Delsuc, F., Kuch, M., Gibb, G.C., Hughes, J., Szpak, P., Southon, J., Enk, J., Duggan, A.T., Poinar, H.N., 2018. Resolving the phylogenetic position of Darwin's extinct ground sloth (Mylodon darwinii) using mitogenomic and nuclear exon data. Proceedings of the Royal Society B: Biological Sciences 285, 20180214.10.1098/rspb.2018.0214. [DOWNLOAD .pdf]

  • Guiry, E., Beglane, F., Szpak, P., Schulting, R., McCormick, F., Richards, M.P., 2018. Anthropogenic changes to the Holocene nitrogen cycle in Ireland. Science Advances 4, eaas9383. doi:10.1126/sciadv.aas9383 [DOWNLOAD .pdf]



  • Szpak, P., Krippner, K., Richards, M.P., 2017. Effects of Sodium Hydroxide Treatment and Ultrafiltration on the Removal of Humic Contaminants from Archaeological Bone. International Journal of Osteoarchaeology 27, 1070-1077. doi:10.1002/oa.2630 [DOWNLOAD .pdf]

  • Szpak, P., Metcalfe, J.Z., Macdonald, R.A. 2017. Best practices for calibrating and reporting stable isotope measurements in archaeology. Journal of Archaeological Science: Reports 13, 609-616.  doi:10.1016/j.jasrep.2017.05.007 [DOWNLOAD .pdf] Appendices: [Appendix A] [Appendix B] [Appendix C] [Appendix D] [Appendix E] [Appendix F] [Appendix G]

  • Braje, T.J., Rick, T.C., Szpak, P., Newsome, S.D., McCain, J.M., Smith, E.A.E., Glassow, M., Hamilton, S.L., 2017. Historical ecology and the conservation of large, hermaphroditic fishes in Pacific Coast kelp forest ecosystems. Science Advances 3, e1601759.  doi:10.1126/sciadv.1601759 [DOWNLOAD .pdf] 

  • Guiry, E., Szpak, P., Richards, M.P., 2017. Isotopic analyses reveal geographical and socioeconomic patterns in historical domestic animal trade between predominantly wheat- and maize-growing agricultural regions in eastern North America. American Antiquity 82, 341-352. doi:10.1017/aaq.2016.34 [DOWNLOAD .pdf] 

  • Carlson, R.L., Szpak, P., Richards, M.P., 2017. The Pender Canal Site and Beginnings of the Northwest Coast Cultural System. Canadian Journal of Archaeology 41, 1-29. [DOWNLOAD.pdf]


  • Szpak, P., Chicoine, D., Millaire, J.-F., White, C.D., Parry, R., Longstaffe, F.J., 2016. Early Horizon camelid management practices in the Nepeña Valley, north-central coast of Peru. Environmental Archaeology 21, 230-245. doi:10.1179/1749631415y.0000000002 [DOWNLOAD .pdf] 

  • Enk, J., Devault, A., Widga, C., Saunders, J., Szpak, P., Southon, J., Rouillard, J.-M., Shapiro, B., Golding, B., Zazula, G., Froese, D., Fisher, D., MacPhee, R., Poinar, H., 2016. Mammuthus Population Dynamics in Late Pleistocene North America: Divergence, Phylogeography and Introgression. Frontiers in Ecology and Evolution 4, 42. doi:10.3389/fevo.2016.00042 [DOWNLOAD .pdf]

  • Guiry, E.J., Needs-Howarth, S., Friedland, K.D., Hawkins, A.L., Szpak, P., Macdonald, R., Courtemanche, M., Holm, E., Richards, M.P., 2016. Lake Ontario salmon (Salmo salar) were not migratory: A long-standing historical debate solved through stable isotope analysis. Scientific Reports 6, 36249. [DOWNLOAD .pdf]

  • Guiry, E.J., Szpak, P., Richards, M.P., 2016. Effects of lipid extraction and ultrafiltration on stable carbon and nitrogen isotopic compositions of fish bone collagen. Rapid Communications in Mass Spectrometry 30, 1591-1600. doi:10.1038/srep36249 [DOWNLOAD .pdf]

  • Jaouen, K., Szpak, P., Richards, M.P., 2016. Zinc Isotope Ratios as Indicators of Diet and Trophic Level in Arctic Marine Mammals. PLoS One 11, e0152299. doi:10.1371/journal.pone.0152299 [DOWNLOAD .pdf]


  • Szpak, P., Millaire, J.-F., White, C.D., Lau, G.F., Surette, F., Longstaffe, F.J., 2015. Origins of Prehispanic Camelid Wool Textiles from the North and Central Coasts of Peru traced by Carbon and Nitrogen Isotopic Analyses. Current Anthropology 56, 449-459. doi:10.1086 /680873 [DOWNLOAD .pdf]

  • Orchard, T.J., Szpak, P., 2015. Zooarchaeological and Isotopic Insights into Locally Variable Economic Patterns: A Case Study from Late Holocene Southern Haida Gwaii, British Columbia. BC Studies 187, 107-147. [DOWNLOAD .pdf]


  • Szpak, P., 2014. Complexities of Nitrogen Isotope Biogeochemistry in Plant-Soil Systems: Implications for the Study of Ancient Agricultural and Animal Management Practices. Frontiers in Plant Science 5, 288. doi:10.3389/fpls.2014.00288 [DOWNLOAD .pdf]

  • Szpak, P., Longstaffe, F.J., Millaire, J.-F., White, C.D., 2014. Large variation in nitrogen isotopic composition of a fertilized legume. Journal of Archaeological Science 45, 72-79.  doi:10.1016/j.jas.2014.02.007 [DOWNLOAD .pdf]

  • Szpak, P., Millaire, J.-F., White, C.D., Longstaffe, F.J., 2014. Small scale camelid husbandry on the north coast of Peru (Virú Valley): Insight from stable isotope analysis. Journal of Anthropological Archaeology 36, 110-129. doi:10.1016/j.jaa.2014.08.005 [DOWNLOAD .pdf]


  • Szpak, P., Orchard, T.J., Salomon, A.K., Gröcke, D.R., 2013. Regional ecological variability and impact of the maritime fur trade on nearshore ecosystems in southern Haida Gwaii (British Columbia, Canada): evidence from stable isotope analysis of rockfish (Sebastes spp.) bone collagen. Archaeological and Anthropological Sciences 5, 159-182. doi:10.1007/s12520-013-0122-y [DOWNLOAD .pdf]

  • Szpak, P., White, C.D., Longstaffe, F.J., Millaire, J.-F., Vásquez Sánchez, V.F., 2013. Carbon and Nitrogen Isotopic Survey of Northern Peruvian Plants: Baselines for Paleodietary and Paleoecological Studies. PLoS One 8, e53763. doi:10.1371/journal.pone.0053763 [DOWNLOAD .pdf]


  • Szpak, P., Longstaffe, F.J., Millaire, J.-F., White, C.D., 2012. Stable Isotope Biogeochemistry of Seabird Guano Fertilization: Results from Growth Chamber Studies with Maize (Zea mays). PLoS One 7, e33741. doi:10.1371/journal.pone.0033741 [DOWNLOAD .pdf]

  • Szpak, P., Millaire, J.-F., White, C.D., Longstaffe, F.J., 2012. Influence of seabird guano and camelid dung fertilization on the nitrogen isotopic composition of field-grown maize (Zea mays). Journal of Archaeological Science 39, 3721-3740. doi:10.1016/j.jas.2012.06.035 [DOWNLOAD .pdf]

  • Szpak, P., Orchard, T.J., McKechnie, I., Gröcke, D.R., 2012. Historical ecology of late Holocene sea otters (Enhydra lutris) from northern British Columbia: isotopic and zooarchaeological perspectives. Journal of Archaeological Science 39, 1553-1571. doi:10.1016/j.jas.2011.12.006 [DOWNLOAD .pdf]


  • Szpak, P., 2011. Fish bone chemistry and ultrastructure: implications for taphonomy and stable isotope analysis. Journal of Archaeological Science 38, 3358-3372. doi:10.1016/j.jas.2011.07.022 [DOWNLOAD .pdf]


  • Szpak, P., Gröcke, D.R., Debruyne, R., MacPhee, R.D.E., Guthrie, R.D., Froese, D., Zazula, G.D., Patterson, W.P., Poinar, H.N., 2010. Regional differences in bone collagen δ13C and δ15N of Pleistocene mammoths: Implications for paleoecology of the mammoth steppe. Palaeogeography, Palaeoclimatology, Palaeoecology 286, 88-96. doi:10.1016/j.palaeo.2009.12.009 [DOWNLOAD .pdf]


  • Szpak, P., Orchard, T.J., Gröcke, D.R., 2009. A Late Holocene vertebrate food web from southern Haida Gwaii (Queen Charlotte Islands, British Columbia). Journal of Archaeological Science 36, 2734-2741. doi:10.1016/j.jas.2009.08.013 [DOWNLOAD .pdf]


  • Debruyne, R., Chu, G., King, C.E., Bos, K., Kuch, M., Schwarz, C., Szpak, P., Gröcke, D.R., Matheus, P., Zazula, G., Guthrie, D., Froese, D., Buigues, B., de Marliave, C., Flemming, C., Poinar, D., Fisher, D., Southon, J., Tikhonov, A.N., MacPhee, R.D.E., Poinar, H.N., 2008. Out of America: ancient DNA evidence for a New World origin of Late Quaternary woolly mammoths. Current Biology 18, 1320-1326. doi:10.1016/j.cub.2008.07.061 [DOWNLOAD .pdf]